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Guppy Color Strains Guppy Color Strains Philip Shaddock Third Edition June 2012 v | Table of Contents Guppy Color Strains © 2000 - 2012 Philip Shaddock his book is copyrighted and may not be reproduced in whole or in part. If you wish to quote more than a few lines from the book or use any of its ig...

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Guppy Color Strains

Guppy Color Strains Philip Shaddock

Third Edition June 2012

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| Table of Contents

Guppy Color Strains © 2000 - 2012 Philip Shaddock his book is copyrighted and may not be reproduced in whole or in part. If you wish to quote more than a few lines from the book or use any of its igures, graphics or images, please contact Philip Shaddock through the Guppy Designer website: www.guppydesigner.com. If you ind inaccuracies or mistakes in the book, please contact Philip Shaddock through the Guppy Designer site. Your help would be very much appreciated. Contact Philip Shaddock at: www.guppydesigner.com 10 9 8 7 6 5 4 3 2 1 ISBN 978-0-9865700-0-1

Guppy Color STrainS

Contents Preface 13 1

Moscows

17

The Moscow in the Rest of the World . . . . . . . . . . . . . . . . . . . . . 18 Moscow Color and Genetics . . . . . . . . . . . . . . . . . . . . . . . . . 19 Breeding the Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . 21 Blue Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23 Albino Blue Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 Blond Blue Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . 24 Asian Blau Blue Moscow . . . . . . . . . . . . . . . . . . . . . . . . . 25 Golden Blue Moscow. . . . . . . . . . . . . . . . . . . . . . . . . . . 26 Green Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 Purple Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27 Full Red Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 28 Half-Black Red Moscow . . . . . . . . . . . . . . . . . . . . . . . . . 29 Albino Full Red Moscow . . . . . . . . . . . . . . . . . . . . . . . . . 30 Golden Red Moscow. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 Midnight Black Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 31 Albino Midnight Black . . . . . . . . . . . . . . . . . . . . . . . . . . 33 Golden Midnight Black . . . . . . . . . . . . . . . . . . . . . . . . . . 34 Half Black Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34 Stoerzbach Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36 Pink White Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 37

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Full Gold Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38 Blue Grass Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 39 Leopard (Grass) Moscow . . . . . . . . . . . . . . . . . . . . . . . . . 40 Carnation Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41 Moscow Fire Tail . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43 WREA Full Pearl Moscow. . . . . . . . . . . . . . . . . . . . . . . . . . . 43 2

Metal Heads

45

Cobra Metal Heads . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 47 Yellow Cobra Metal Head . . . . . . . . . . . . . . . . . . . . . . . . . 47 Red Cobra Metal Head Roundtail . . . . . . . . . . . . . . . . . . . . . 47 Black Cobra Metal Head . . . . . . . . . . . . . . . . . . . . . . . . . 48 Red Metal Head Mosaic . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49 Lace Metal Heads . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 50 Red Lace Metal Head. . . . . . . . . . . . . . . . . . . . . . . . . . . 50 Orange Metal Head Fantail . . . . . . . . . . . . . . . . . . . . . . . . 50 Yellow Lace Metal Head Fantail . . . . . . . . . . . . . . . . . . . . . . 51 Yellow Metal Head Doublesword . . . . . . . . . . . . . . . . . . . . . 51 Yellow Lace Metal Head Speartail . . . . . . . . . . . . . . . . . . . . . 52 3

Snakeskins

53

Common Names. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54 Snakeskin Guppy History . . . . . . . . . . . . . . . . . . . . . . . . . 54 The Pattern. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 58 Snakeskin Variations. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 59 The Lace Snakeskin Genetics . . . . . . . . . . . . . . . . . . . . . . . 64 Zebrinus or Bar Genetics . . . . . . . . . . . . . . . . . . . . . . . . . 64 Yellow Cobra Snakeskin Roundtail . . . . . . . . . . . . . . . . . . . . . . 67 Albino Yellow Cobra Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . 68

Guppy Color STrainS

Blond Snakeskin Speartail . . . . . . . . . . . . . . . . . . . . . . . . . . 69 Black Snakeskin. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69 Speartail Snakeskin Saddleback . . . . . . . . . . . . . . . . . . . . . . . 70 Yellow Cobra Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . . 70 Cobra Snakeskin Doublesword . . . . . . . . . . . . . . . . . . . . . . . . 71 Lace Snakeskin Upper Sword. . . . . . . . . . . . . . . . . . . . . . . . . 72 Green Lace Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73 Yellow Mosaic Snakeskin. . . . . . . . . . . . . . . . . . . . . . . . . . . 74 Purple Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74 Blue Half-Moon Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . 75 Red Lace Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 75 Glass Belly Red Lace Snakeskin Swallow . . . . . . . . . . . . . . . . . . . . 77 Ribbon Red Roundtail . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77 Red Snakeskin Roundtail. . . . . . . . . . . . . . . . . . . . . . . . . . . 78 Full Gold Snakeskin . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78 4

Magentas

81

Long Fin Silverado. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83 Short Fin Silverado . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 83 Blue Silverado. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 84 Full Gold Magenta Moscow. . . . . . . . . . . . . . . . . . . . . . . . . . 85 Blond Magenta Moscow . . . . . . . . . . . . . . . . . . . . . . . . . . . 86 Albino Full Red Magenta (Pink) . . . . . . . . . . . . . . . . . . . . . . . . 87 Albino Metallic Magenta Ribbon . . . . . . . . . . . . . . . . . . . . . 87 Flamenco Dancer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 88 Peacock . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 89 Magenta Metal Lace Cobra . . . . . . . . . . . . . . . . . . . . . . . . . . 90 5

Medusas and Galaxies

91

Medusa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 93

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Medusa Roundtail . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94 Platinum Medusa . . . . . . . . . . . . . . . . . . . . . . . . . . . . 95 European Spadetail Medusa . . . . . . . . . . . . . . . . . . . . . . . 96 Blue Galaxy. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 97 Red Galaxy Fantail . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 White Galaxy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 98 European Magenta Galaxy . . . . . . . . . . . . . . . . . . . . . . . . 99 Yellow Grass Galaxy . . . . . . . . . . . . . . . . . . . . . . . . . . 100 Blue Grass Galaxy . . . . . . . . . . . . . . . . . . . . . . . . . . . 100 Santa Maria . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 101 Albino Red Santa Maria . . . . . . . . . . . . . . . . . . . . . . . . 103 6

Grass

105

Blue Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 107 Red Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 109 Albino Red Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . 109 Albino Full Red Grass. . . . . . . . . . . . . . . . . . . . . . . . . . 110 Yellow Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 110 Purple Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 111 Silver Grass . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 111 7

Full Golds and Full Platinums

113

The White Saddle Gene. . . . . . . . . . . . . . . . . . . . . . . . . 115 Platinum Versus Gold Metal . . . . . . . . . . . . . . . . . . . . . . 115 White Full Gold Leucophore . . . . . . . . . . . . . . . . . . . . . . 118 Gray Full Platinum . . . . . . . . . . . . . . . . . . . . . . . . . . . 118 Full Platinum Redtail . . . . . . . . . . . . . . . . . . . . . . . . . 119 Albino Blue Platinum . . . . . . . . . . . . . . . . . . . . . . . . . . . 120 Albino Full Gold Blau . . . . . . . . . . . . . . . . . . . . . . . . . . . 121

Guppy Color STrainS

8

Schimmelpfennig Platinums

123

Schimmelpfennig Platinum Sword . . . . . . . . . . . . . . . . . . . . . 125 Double Yellow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128 El Dorado. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 129 Albino El Dorado . . . . . . . . . . . . . . . . . . . . . . . . . . . 130 Red Fin Metal Speartail . . . . . . . . . . . . . . . . . . . . . . . . . . 132 Blond Platinum Speartail . . . . . . . . . . . . . . . . . . . . . . . . . 132 Platinum Red Mosaic . . . . . . . . . . . . . . . . . . . . . . . . . . . 134 9

Pink Whites

137

Snow White . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 138 Green Platinum Pink White. . . . . . . . . . . . . . . . . . . . . . . . . 139 Albino Japanese Pink White Swallow . . . . . . . . . . . . . . . . . . . . 140 10

Pinks and Pingus

141

Pingu . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 141 Pingu Panda . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 143 Platinum Pingu . . . . . . . . . . . . . . . . . . . . . . . . . . . . 144 Panda Moscow (Pink Moscow) . . . . . . . . . . . . . . . . . . . . . . . 145 Glass Belly Panda . . . . . . . . . . . . . . . . . . . . . . . . . . . 146 Blond Glass Belly Panda . . . . . . . . . . . . . . . . . . . . . . . . 147 11

Blue Metal Guppies

149

Blue Diamond (Luster) . . . . . . . . . . . . . . . . . . . . . . . . . . . 149 Full Metal Ivory Blue Mosaic Full Moon . . . . . . . . . . . . . . . . . . . 150 Lazuli . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 Lace Snake Lazuli . . . . . . . . . . . . . . . . . . . . . . . . . . . 153 Turquoise . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 153 Japan Blue (Aquamarine) . . . . . . . . . . . . . . . . . . . . . . . . . 155 Albino Japan Blue . . . . . . . . . . . . . . . . . . . . . . . . . . . 157

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Wild Japan Blue . . . . . . . . . . . . . . . . . . . . . . . . . . . . 157 Blue Metal Doublesword. . . . . . . . . . . . . . . . . . . . . . . . . . 158 12

Stoerzbach

159

Stoerzbach Metal Swordtail . . . . . . . . . . . . . . . . . . . . . . . . 159 Pied Blue Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 163 13

Micarifs

165

Orange Micarif . . . . . . . . . . . . . . . . . . . . . . . . . . . . 168 14

Red Guppies

169

Red Pigments. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 171 Full Red . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 173 Blond Half-Black Full Red . . . . . . . . . . . . . . . . . . . . . . . . . 176 Albino Full Red . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 177 Albino Full Red Swallow . . . . . . . . . . . . . . . . . . . . . . . . 178 Marlboro Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 178 Albino Glass Belly Red Speartail . . . . . . . . . . . . . . . . . . . . . . 179 Mosaic Red Fantail . . . . . . . . . . . . . . . . . . . . . . . . . . . . 180 Blond Big Dorsal Red Tail. . . . . . . . . . . . . . . . . . . . . . . . . . 181 Coral Red Doublesword . . . . . . . . . . . . . . . . . . . . . . . . . . 181 Platinum Firetail Full Moon . . . . . . . . . . . . . . . . . . . . . . . . 182 15

Half-Black or Tuxedo

185

Evidence For The Two Gene Theory . . . . . . . . . . . . . . . . . . . . . 187 Area of Coverage . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 187 Half-Black Dominance . . . . . . . . . . . . . . . . . . . . . . . . . 189 Breeding Half-Blacks . . . . . . . . . . . . . . . . . . . . . . . . . . . 190 Half-Black Yellow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 191 Half-Black Red . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 193

Guppy Color STrainS

Glass Belly Half-Black Red . . . . . . . . . . . . . . . . . . . . . . . 194 Half-Black Purple . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 195 Half-Black Pastel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 197 Blond Half-Black Pastel (White and Yellow Pastel) . . . . . . . . . . . . 199 Half-Black White . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 200 RREA Yellow Tuxedo . . . . . . . . . . . . . . . . . . . . . . . . . . 201 Half-Black Green . . . . . . . . . . . . . . . . . . . . . . . . . . . 202 Black Half-Black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 203 Dragon Head or Leopard . . . . . . . . . . . . . . . . . . . . . . . . . . 204 16

Solid Color

207

American Blue Delta. . . . . . . . . . . . . . . . . . . . . . . . . . . . 207 American Green Delta . . . . . . . . . . . . . . . . . . . . . . . . . . . 208 American Purple Delta . . . . . . . . . . . . . . . . . . . . . . . . . . . 211 American Bronze (Golden) . . . . . . . . . . . . . . . . . . . . . . . . . 212 17

Colorless Strains

213

Albino Blau. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 213 Super White . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 214 Ruby Eye Super White (WREA Super White) . . . . . . . . . . . . . . . 216 See-thru . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 216 Blond See-thru . . . . . . . . . . . . . . . . . . . . . . . . . . . . 218 18

Variegated, Wild and Endler

221

Rio Leao Feral Guppies . . . . . . . . . . . . . . . . . . . . . . . . . . . 221 Erfurt Wild Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222 Lampang Wild Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . 223 Istanbul . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 223 Vienna Emerald Green Swordtail . . . . . . . . . . . . . . . . . . . . . . 224 Single Sword Vienna Emerald Green . . . . . . . . . . . . . . . . . . . 226

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Old Fashioned . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 227 Top Yellow Sword 2004 Endler . . . . . . . . . . . . . . . . . . . . . . . 228 Tangerine Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 230 White Tangerine. . . . . . . . . . . . . . . . . . . . . . . . . . . . 231 Balloon Guppy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231 A

Appendix: Gene Symbols

235

Sex-Linked Notation . . . . . . . . . . . . . . . . . . . . . . . . . . . 236 B

Appendix: Gene Table

237

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Preface his book has been twelve years in the making and has had previous incarnations. It now includes over 150 strains. It’s accuracy, in terms of correctly identifying the genes involved in each of the strains, has been steadily improving because I have done many of the crosses myself. My goal throughout the last twelve years is to provide guppy breeders with a reliable reference source. Some will use the collection to identify guppies they acquired but whose genetics they do not know. Others will use the book to ind out how to breed speciic strains. Some strains can be fairly complex and require basic knowledge of their genotype to maintain them. Still others will use the book to ind strains they are interested in breeding. he guppy breeder interested in crossing experiments will ind that many combinations of color genes have already been created and reined. I have grouped many variants of basic strains together to make the collection useful. For example, you can see what a White HalfBlack Pastel looks like, then the blond white and yellow variants. I have also provided a lot of breeding tips for speciic strains, including tips for developing speciic strains for the judging bench. Where the gene makeup of the strain is complex I provide basic information about how to keep all those genes intact. Classiication System here is no good way to absolutely categorize guppies into strains, where a strain is deined as a truebreeding guppy that produces sons and daughters that are 80% similar. An example is a Panda Moscow, a guppy with both Moscow with the Pink mutations (Fig.1). he Panda belongs to both Moscow and Pink categories. In loosely grouping guppies in this book, I start of with a rule that says certain strains have dominant traits that are strictly Y-linked.

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he Moscow is a case in point. he Moscow deining characteristics are strongly Y-linked, meaning they are passed on from father to son and not to daughters. So I have grouped all guppies that have the Moscow Y-linked supergene together as a family of strains.

Fig.1 Panda Moscow. Photo by Finn Bindeballe

hen there is the “Blond Glass Belly Panda,” which combines the blond, Glass Belly, Pink and Moscow mutations (Fig.2 ). his is a guppy that belongs to three diferent categories. I know it is entirely arbitrary, but I think of the Blond and Glass Belly traits as “modiier” traits because they are autosomal and tend to afect the whole body uniformly. So I have made the Blond Glass Belly Panda a variant of the Panda and the Panda is grouped with the Pink mutation guppies. he Color Strains book has split many of the old groupings into smaller groupings. For example, the Mg (Metallic Gold) gene has now got its own chapter. I have nothing to recommend my arbitrary classiications other than the fact that they may be of some use to breeders examining strains of unknown genotype. Or more simply the classiication system makes this book easier to read from beginning to end. In fact fancy guppies are usually an arbitrary concoction of genes, selected for aesthetic appeal and perpetuated through the habit of selective breeding and the assignment of such names as “Flamenco Dancer.” here are so-called “base body” classiication systems. Classiication

Guppy Color STrainS

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systems like these have to do with phenotype, what you see. My work with the microscope examining color on the cellular level shows that what you think you see with your naked eye is not necessarily what you actually see with the aid of the microscope. he microscope is much more accurate. And if you had tools that could make observations on the molecular level, your observations would be that much more accurate again. Good theories rest on accurate observations. he designer is interested in genotype, the actual genes that are determining the appearance of the guppy.

Fig. 2 Glass Belly Panda. Photo: Philip Shaddock

I record the genotype for many of the strains in this book. Ultimately this is the most accurate way to identify guppies. You will ind a guide to the system of notation I use for genotype in Appendix A: Gene Symbols. I recommend that you buy he heory and Practice of Guppy Breeding as a companion to this book. It will make understanding the genetics sections a lot easier. And I include a wide array of breeding strategies and advice on designing breeding programs. Neither book stands on its own.

Acknowl...